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How is Desmond Physically Fit?

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He spends most of his time strapped in an animus yet he seems strong and agile enough for combat and parkour. I understand the animus teaching Desmond HOW to use Assassin free running and fighting techniques; but how is he physically able to carry them out with complete precision?

Does the animus transfer physical capabilities? I think not because Lucy once remarked how he was getting fat. So how could he then climb the Colosseum, infiltrate abstergo etc.?

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